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Table 4 Species expanding their distribution into new MSFD areas

From: Mediterranean non indigenous species at the start of the 2020s: recent changes

Higher taxon Species From To status pathway Expansion record(s)
ASC Perophora multiclathrata (Sluiter, 1904) WMED CMED unk T-S (fouling) IT: Lezzi et al., 2018
PHY Chaetoceros bacteriastroides G.H.H.Karsten, 1907 WMED ADRIA est T-S (ballast) HR: Čalić, M., et al., 2018
BRY Parasmittina egyptiaca (Waters, 1909) EMED CMED unk T-S (fouling) IT: Ulman et al., 2017
BRY Watersipora arcuata Banta, 1969 WMED CMED, EMED est T-S (fouling) ES, IT, MT, TR: Ulman et al., 2017
CHL Ulva ohnoi M.Hiraoka & S.Shimada, 2004 WMED CMED, EMED est T-S (fouling) IL: Krupnik et al., 2018 / TN: Miladi et al., 2018
CRU Paracaprella pusilla Mayer, 1890 EMED, WMED CMED est T-S (fouling) TN: Fersi et al., 2018
CRU Stenothoe georgiana Bynum & Fox, 1977 WMED CMED est T-S (fouling) IT: Ulman et al., 2017
CRU Oithona davisae Ferrari F.D. & Orsi, 1984 WMED EMED, CMED, ADRIA est T-S (ballast) GR: Dragičević et al., 2019; TR: Terbıyık Kurt & Beşiktepe, 2019 / IT: Zagami et al., 2018 / IT: Vidjak et al., 2019
CRU Dyspanopeus sayi (Smith, 1869) CMED EMED cas T-S (ballast)/CON GR: Ulman et al., 2017
CRU Anilocra pilchardi Bariche & Trilles, 2006 EMED, CMED WMED unk UNA TN: Ounifi Ben Amor et al., 2017
CRU Ianiropsis serricaudis Gurjanova, 1936 ADRIA WMED est T-S (fouling) IT: Ferrario et al., 2017 / FR: Ulman et al., 2017
FISH Bregmaceros nectabanus Whitley 1941 EMED CMED est UNA GR: Ketsilis-Rinis & Dimitriou in Chartosia et al., 2018
FISH Etrumeus golanii Di Battista, Randall & Bowen, 2012 EMED, CMED WMED est UNA DZ: Kassar & Hemida in Stamouli et al., 2017 / TN: Rafrafi-Nouira et al., 2017 / MA: Tamsouri et al., 2019
FISH Parupeneus forsskali (Fourmanoir & Guézé, 1976) EMED CMED est UNA TN: Capapé et al., 2018b
FISH Pteragogus trispilus Randall, 2013 EMED, CMED WMED cas UNA TN: Hamed et al., 2018
FISH Pterois miles (Bennett, 1828) EMED, WMED CMED est UNA IT: Azzurro et al., 2017 / GR: Mitsou & Maximiadi in Yokeş et al., 2018 / LY: Al Mabruk & Rizgalla, 2019
FISH Sciaenops ocellatus (Linnaeus, 1766) EMED CMED est UNA IT: Langeneck et al., 2017
FISH Synchiropus sechellensis Regan, 1908 GR: EMED CMED unk UNA GR: Teneketzis & Christidis in Yokeş et al., 2018
MOL Dendostrea cf folium (Linnaeus, 1758) EMED ADRIA est T-S (fouling) AL: Xharahi et al. in Gerovasileiou et al., 2017
MOL Fulvia fragilis (Forsskål in Niebuhr, 1775) EMED, CMED, WMED ADRIA est T-S/UNA AL: Xharahi et al. in Gerovasileiou et al., 2017 / ME: Gvozdenović et al., 2019
MOL Malleus regula (Forsskål in Niebuhr, 1775) EMED CMED est T-S (fouling) IT: Prato & Rubino in Kousteni et al., 2019
MOL Sepioteuthis lessoniana Férussac [in Lesson], 1831 EMED, CMED ADRIA est UNA ME: Dragičević et al. in Stern et al., 2019
MOL Biuve fulvipunctata (Baba, 1938) EMED, CMED, WMED ADRIA est UNA HR: Petani & Crocetta in Kousteni et al., 2019
MOL Conomurex persicus (Swainson, 1821) EMED, CMED ADRIA unk T-S/UNA AL: Xharahi et al. in Gerovasileiou et al., 2017
MOL Diodora ruppellii (G. B. Sowerby I, 1835) EMED, WMED CMED cas? T-S (fouling)/UNA LY: Rizgalla et al., 2019
MOL Godiva quadricolor (Barnard, 1927) WMED, ADRIA CMED est T-S/CON IT: Furfaro et al., 2018
MOL Haminoea cyanomarginata Heller & Thompson, 1983 EMED, CMED, WMED ADRIA unk T-S/UNA HR: Dragicevic et al. in Chartosia et al., 2018
PLAT Allolepidapedon fistulariae Yamaguti, 1940 EMED, WMED CMED unk UNA LY: Salem, 2017
PLAT Glyphidohaptor plectocirra (Paperna, 1972) Kritsky, Galli & Yang, 2007 EMED CMED unk UNA LY: Abdelnor et al., 2019
PLAT Maritigrella fuscopunctata (Prudhoe, 1978) CMED EMED unk T-S/UNA IL: Velasquez et al., 2018
POL Chaetozone corona Berkeley & Berkeley, 1941 EMED, CMED ADRIA, WMED est T-S (ballast) IT (ADRIA): Munari et al., 2017; Grossi et al., 2017 / IT (WMED): Munari et al., 2017
POL Desdemona ornata Banse, 1957 EMED, ADRIA, WMED CMED unk T-S/CON TN: Boudaya et al., 2019
POL Dorvillea similis (Crossland, 1924) EMED WMED est T-S (fouling)/UNA IT: Langeneck & Tempesti in Dragicevic et al., 2019
POL Hydroides brachyacantha Rioja, 1941 EMED WMED unk T-S (fouling) ES: Ulman et al., 2017
POL Palola valida (Gravier, 1900) EMED ADRIA unk T-S (fouling) ME: Spagnolo et al., 2019
RHO Chondria curvilineata F.S.Collins & Hervey, 1917 EMED, WMED CMED est T-S IT: Alongi in Gerovasileiou et al., 2017
  1. The pathway column refers to the most likely pathway of secondary spread. Taxonomic group and Pathway abbreviations as in Table 2